To my eye, the Tempest's design features fix several of the short-comings of existing LO/VLO fighter aircraft:
1. The explicit statement of "balanced-survivability" is a rejection, either through operational analysis or budget analysis, of the pursuit of even greater stealth. I suspect operational analysis, as we are starting to see more and more comments to the effect that stealth is a necessary, but not sufficient, component of future aircraft survivability.
2. The reconfigurable payload bay and planned conformal carry is an explicit attempt to return some flexibility to stealth aircraft. I suspect the origins are that the F-22 and F-35 do not have a lot of room internally to fit different payloads. For the USAF, this is less of an issue because the USAF can afford other support aircraft to carry payloads. For the RAF and like-minded airforces, this will be an issue. China and Russia have already moved towards excessively large bays on their stealth aircraft, this is a continuation and amplification of that trend.
Now, the curious question is if we will see similar design decisions made in the PCA. I think so.